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Revision Of The Mesobuthus Caucasicus Complex From Central Asia, With Descriptions Of Six New Species (Scorpiones: Buthidae), Victor Fet, František Kovařík, Benjamin Gantenbein, Ronald C. Kaiser, Alexander K. Stewart, Matthew R. Graham Mar 2018

Revision Of The Mesobuthus Caucasicus Complex From Central Asia, With Descriptions Of Six New Species (Scorpiones: Buthidae), Victor Fet, František Kovařík, Benjamin Gantenbein, Ronald C. Kaiser, Alexander K. Stewart, Matthew R. Graham

Victor Fet

A widespread Mesobuthus caucasicus complex, which includes some of the most common scorpions found from the Caucasus to China, is revised for the first time based on new extensive collections from Central Asia, using both morphological and DNA marker data. Mesobuthus caucasicus (Nordmann, 1840), s.str. is restricted to the Caucasus Mts. Four taxa are elevated to species rank: M. fuscus (Birula, 1897) (Tajikistan), M. intermedius (Birula, 1897) (Tajikistan), M. kaznakovi (Birula, 1904) (Tajikistan, Uzbekistan), and M. parthorum (Pocock, 1889) (Afghanistan, Iran, Turkmenistan). Six new species are described: M. brutus sp. n. (Iran), M. elenae sp. n. (Tajikistan, Uzbekistan), M ...


Catalinia, A New Scorpion Genus From Southern California, Usa And Northern Baja California, Mexico (Scorpiones: Vaejovidae), Michael E. Soleglad, Richard F. Ayrey, Matthew R. Graham, Victor Fet Nov 2017

Catalinia, A New Scorpion Genus From Southern California, Usa And Northern Baja California, Mexico (Scorpiones: Vaejovidae), Michael E. Soleglad, Richard F. Ayrey, Matthew R. Graham, Victor Fet

Victor Fet

Genus Catalinia, gen. nov. (Scorpiones: Vaejovidae) is described from southern California, USA and Baja California, Mexico. The genus is composed of four species formerly placed in Pseudouroctonus: Catalinia minima (Kraepelin, 1911), comb. nov. (type species), C. andreas (Gertsch et Soleglad, 1972), comb. nov., C. castanea (Gertsch et Soleglad, 1972), comb. nov., and C. thompsoni, comb. nov. (Gertsch et Soleglad, 1972). Major diagnostic characters of Catalinia include a carapace with a very weak anterior indentation, a very stout metasoma with little or no tapering from segment I to V, and a mating plug with two partial bases. Evidence is presented suggesting ...


The Second Record Of A Relict Akrav Israchanani Levy, 2007 (Scorpiones: Akravidae) From Levana Cave, Israel, Victor Fet, Michael E. Soleglad, Sergei L. Zonstein, Israel Naaman, Shlomi Lubaton, Boaz Langford, Amos Frumkin Sep 2017

The Second Record Of A Relict Akrav Israchanani Levy, 2007 (Scorpiones: Akravidae) From Levana Cave, Israel, Victor Fet, Michael E. Soleglad, Sergei L. Zonstein, Israel Naaman, Shlomi Lubaton, Boaz Langford, Amos Frumkin

Victor Fet

We report the remnants of five new scorpion specimens discovered dead in Levana Cave in Israel in December 2015. We confirm that they belong to the relict scorpion Akrav israchanani Levy, 2007 (Akravidae), famously described from the neighboring Ayyalon Cave, also from dead specimens. The details of morphology of the new specimens are given; they match completely the characters of A. israchanani redescribed by Fet, Soleglad & Zonstein (2011). This second record indicates a wider distribution of this unique cave scorpion, which, however, is extinct in both caves. There is still no evidence that live populations of this species exist.


Two New Euscorpius Species From Central-Western Greece (Scorpiones: Euscorpiidae), Gioele Tropea, Victor Fet Dec 2016

Two New Euscorpius Species From Central-Western Greece (Scorpiones: Euscorpiidae), Gioele Tropea, Victor Fet

Victor Fet

Two new Euscorpius species are described, based on specimens collected by P.M. Giachino & D. Vailati in central-western Greece in neighboring Aitoloakarnania (Western Greece) and Fokida (Central Greece) regional units. No Euscorpius specimens were previously available from this area. The first new species, Euscorpius giachinoi sp. n., is very similar to a recently described E. birulai Fet et al., 2014 from Euboea Island, and is also characterized by a low trichobothrial count (Pv = 7, et = 5), a low pectinal teeth count (Dp = 7 in males, 6 in females), and long-limbed features. The second species, E. vailatii sp. n., is widely found in the studied area, and it is characterized by a high trichobothrial and pectinal teeth count (Pv = 9–11, et = 6–8; Dp = 9–10 in males, 7–8 in females).


Three New Species Of Euscorpius (Scorpiones: Euscorpiidae) From Greece, Gioele Tropea, Victor Fet, Aristeidis Parmakelis, Panayiota Kotsakiozi, Iasmi Stathi Dec 2016

Three New Species Of Euscorpius (Scorpiones: Euscorpiidae) From Greece, Gioele Tropea, Victor Fet, Aristeidis Parmakelis, Panayiota Kotsakiozi, Iasmi Stathi

Victor Fet

Three new species of the genus Euscorpius Thorell, 1876 are described from Greece: E. stahlavskyi sp.n. from Epirus in the northwestern Greece; E. kinzelbachi sp.n. from Mt. Olympus at the eastern border between Thessaly and Central Macedonia, and E. vignai sp.n. from Karpathos and Kasos Islands (eastern Aegean Sea). Species-level divergence of these taxa is also confirmed by multiple DNA markers in Parmakelis et al. (2013).


The Systematic Position Of The Scorpion Genera Heteroscorpion Birula, 1903 And Urodacus Peters, 1861 (Scorpiones: Scorpionoidea), Michael E. Soleglad, Victor Fet, František Kovařík Dec 2016

The Systematic Position Of The Scorpion Genera Heteroscorpion Birula, 1903 And Urodacus Peters, 1861 (Scorpiones: Scorpionoidea), Michael E. Soleglad, Victor Fet, František Kovařík

Victor Fet

Stockwell (1989), in an unpublished revision, suggested the elevation of subfamily Urodacinae to the family rank, and considered Heteroscorpioninae a subfamily of Ischnuridae. Heteroscorpioninae was formally elevated to the family rank by Lourenço (1996a), and Urodacinae, by Prendini (2000). As a result of a detailed cladistic analysis, Prendini (2000, 2003b) considered families Heteroscorpionidae and Urodacidae to be sister groups, although an al-ternative topology was available. Soleglad & Fet (2003b) questioned the results of Prendini (2000) but nevertheless retained the monophyly of Heteroscorpion and Urodacus pending more detailed analysis; they recognized two valid monotypic subfamilies, Heteroscorpioninae and Urodacinae, under Urodacidae. Our present detailed cladistic re-analysis of Heteroscorpion and Urodacus confirms the phylogeny proposed by Stockwell (1989), which also is the “suboptimal” topology of ...


Vestigial Serrula In Scorpion Genera Paravaejovis, , Smeringurus And Vejovoidus (Scorpiones: Vaejovidae), Victor Fet, Michael E. Soleglad, Michael S. Brewer, David P. A. Neff, Michael L. Norton Dec 2016

Vestigial Serrula In Scorpion Genera Paravaejovis, , Smeringurus And Vejovoidus (Scorpiones: Vaejovidae), Victor Fet, Michael E. Soleglad, Michael S. Brewer, David P. A. Neff, Michael L. Norton

Victor Fet

The presence of vestigial serrula on the ventral edge of the cheliceral movable finger is established for vaejovid genera Paravaejovis, Paruroctonus, Smeringurus, and Vejovoidus. Detailed descriptions and SEM images illustrating the various manifestations of this interesting cheliceral structure are provided for many species of these four genera. A brief systematic overview of this structure as it exists in Recent scorpions is also presented.


Three More Species Of Euscorpius Confirmed For Greece (Scorpiones: Euscorpiidae), Victor Fet, Michael E. Soleglad, Aristeidis Parmakelis, Panayiota Kotsakiozi, Iasmi Stathi Dec 2016

Three More Species Of Euscorpius Confirmed For Greece (Scorpiones: Euscorpiidae), Victor Fet, Michael E. Soleglad, Aristeidis Parmakelis, Panayiota Kotsakiozi, Iasmi Stathi

Victor Fet

In Greece, scorpion genus Euscorpius has been insufficiently studied. Taxonomy of several species and subspecies has been inconsistent and confusing. We provide new morphological data and redescriptions of type specimens of three “old” taxa, described from Greece and formerly listed under a “catch-all” taxon Euscorpius carpathicus. We elevate to, or confirm at species status: Euscorpius scaber Birula, 1900 (type locality: Mt. Athos), E. candiota Birula, 1903 (type locality: Heraklion, Crete), and E. ossae Di Caporiacco, 1950, stat.n. (type locality: Mt. Ossa, Thessaly). Species-level separation of these taxa is also confirmed by multiple species delimitation methods implemented on the phylogenetic ...


The First Record Of Upper Permian And Lower Triassic Scorpions From Russia (Chelicerata: Scorpiones), Victor Fet, Dmitry E. Shcherbakov, Michael E. Soleglad Dec 2016

The First Record Of Upper Permian And Lower Triassic Scorpions From Russia (Chelicerata: Scorpiones), Victor Fet, Dmitry E. Shcherbakov, Michael E. Soleglad

Victor Fet

Several small fragments of fossil scorpions are reported from two localities in Vologda Province, Russia, representing the Upper Permian (Severodvinian, correlated to Wuchiapingian) (Isady) and Lower Triassic just above the Permian-Triassic boundary (Induan) (Nedubrovo). Most observed structures are not diagnostic at genus or family level. The Isady leg fragment possesses ungues (claws), which are both denticulated and setaceous, and resembles a Carboniferous Eobuthus sp. (Eobuthidae). It is the latest record of this type of ungues, which are known in some Paleozoic scorpions (extinct suborder Mesoscorpiones); all extant scorpions have smooth claws without denticulation or setation.


The Genus Akrav Levy, 2007 (Scorpiones: Akravidae) Revisited, Victor Fet, Michael E. Soleglad, Sergei L. Zonstein Dec 2016

The Genus Akrav Levy, 2007 (Scorpiones: Akravidae) Revisited, Victor Fet, Michael E. Soleglad, Sergei L. Zonstein

Victor Fet

Akrav israchanani, a relict chactoid scorpion from the famous Ayyalon Cave in Israel, is analyzed for the first time since its original description by Gershom Levy (2007). All scorpions found in this cave (20 specimens) were dead, represented by exoskeletons; they are mostly fragmented during collection, many incomplete, but extremely well preserved, and have no evidence of fossilization. Time and cause of death are unknown. Diagnostic characters described by Levy are largely confirmed, and some are further clarified. An exhaustive set of microscopic images is published, encompassing data from all best preserved specimens. Previously unpublished morphological details are illustrated such ...


Taxonomic Position Of The Genus Simonoides Vachon Et Farzanpay, 1987, And Description Of A New Species Of Orthochirus Karsch From Iran (Scorpiones, Buthidae), František Kovařík, Victor Fet Dec 2016

Taxonomic Position Of The Genus Simonoides Vachon Et Farzanpay, 1987, And Description Of A New Species Of Orthochirus Karsch From Iran (Scorpiones, Buthidae), František Kovařík, Victor Fet

Victor Fet

Simonoides Vachon et Farzanpay, 1987 is synonymized with Orthochirus Karsch, 1891. Lectotype is designated for Simonoides farzanpayi Vachon et Farzanpay, 1987. Orthochirus sobotniki Kovařík, 2004 is synonymized with Orthochirus farzanpayi (Vachon et Farzanpay, 1987), comb. n. Orthochirus gruberi, sp. n. (Iran) is described and dis-tinguished from all other Iranian species of the genus on a combination of two characters, the presence of rows of granules with internal and external granules on the movable fingers of pedipalps and hirsuteness of the entire me-tasoma and telson.


The Scorpion Sternum: Structure And Phylogeny (Scorpiones: Orthosterni), Michael E. Soleglad, Victor Fet Dec 2016

The Scorpion Sternum: Structure And Phylogeny (Scorpiones: Orthosterni), Michael E. Soleglad, Victor Fet

Victor Fet

The structure of the sternum of all major Recent scorpion groups is analyzed in detail. Based on this analysis, two fundamental sternum types are identified, described and illustrated, type 1 and type 2. These sternum types are distinguished by criteria based on external and internal structural features. The sternum types described herein are offered as a replacement for the various characterizations used throughout the last 140 years which emphasize only gross overall shape and proportions. Phylogenetic and taxonomic ramifications of these new sternal types are discussed. The Carboniferous fossil scorpion Palaeopisthacanthus schucherti Petrunkevitch is assigned to sternum type 1. The ...


The Genus Auyantepuia González-Sponga, 1978 Is Confirmed As A Synonym Of Broteochactas Pocock, 1893 (Scorpiones: Chactidae), Michael E. Soleglad, Victor Fet Dec 2016

The Genus Auyantepuia González-Sponga, 1978 Is Confirmed As A Synonym Of Broteochactas Pocock, 1893 (Scorpiones: Chactidae), Michael E. Soleglad, Victor Fet

Victor Fet

The genus Auyantepuia González-Sponga, 1978 is confirmed as a synonym of Broteochactas Pocock, 1893 (Scorpiones: Chactidae). It was correctly synonymized with Broteochactas by Francke & Boos (1986) and listed as such by Soleglad & Fet (2003) but resurrected by Lourenço & Araújo (2004). We reconfirm that the generic name Auyantepuia, based on the type species Broteochactas scorzai Dagert, 1957 from Venezuela, is a synonym of Broteochactas, based on the type species Broteochactas nitidus Pocock, 1893 from Trinidad. All other seven “short-fingered” species from French Guiana and Brazil, classified by Lourenço & Araújo (2004) under Auyantepuia, belong to the genus Neochactas Soleglad et Fet, 2003 ...


The First Record Of The Genus Anomalobuthus Kraepelin, 1900 From Iran, With Description Of A New Species (Scorpiones: Buthidae), Rolando Teruel, František Kovařík, Shahrokh Navidpour, Victor Fet Dec 2016

The First Record Of The Genus Anomalobuthus Kraepelin, 1900 From Iran, With Description Of A New Species (Scorpiones: Buthidae), Rolando Teruel, František Kovařík, Shahrokh Navidpour, Victor Fet

Victor Fet

We describe herein a new species of buthid scorpions: Anomalobuthus talebii sp. n. This is the second species assigned to this genus, endemic from sand deserts of Central Asia, as well as the first record of this genus from Iran. The new species is fully illustrated with color photos of holotype habitus, key morphologically diagnostic characters, and habitat.


Serrula In Retrospect: A Historical Look At Scorpion Literature (Scorpiones: Orthosterni), Matthew R. Graham, Victor Fet Dec 2016

Serrula In Retrospect: A Historical Look At Scorpion Literature (Scorpiones: Orthosterni), Matthew R. Graham, Victor Fet

Victor Fet

Serrula, a small comb-like structure on the ventral aspect of cheliceral movable finger in scorpions, has often been overlooked or ignored. Scorpion genera that are currently known to possess serrula belong to six families: Chactidae, Euscorpiidae, Iuridae, Pseudochactidae, Superstitioniidae, and Vaejovidae. All relevant literature on scorpion serrula is reviewed, and serrula of representative scorpion taxa is illustrated by SEM images. We suggest that the inconsistency in reporting serrula is based on several factors (or combination thereof): (a) serrula tines can quite often be broken off partially or completely, especially in adults; (b) serrula can be vestigial and hardly visible under ...


Taxonomic Position Of The Genus Sassanidotus Farzanpay, 1987 (Scorpiones: Buthidae), František Kovařík, Victor Fet Dec 2016

Taxonomic Position Of The Genus Sassanidotus Farzanpay, 1987 (Scorpiones: Buthidae), František Kovařík, Victor Fet

Victor Fet

Sassanidotus Farzanpay, 1987 is a valid genus closely related to Compsobuthus Vachon, 1949, with which it shares most characters including the shape of carinae on the carapace and mesosoma. The main difference is that Sassanidotus has only three granules located proximally to the terminal granule on the movable finger of pedipalp, whereas Compsobuthus and also Mesobuthus Vachon, 1950 have four such granules. Sassanidotus Farzanpay, 1987 includes two species, Sassanidotus zarudnyi (Birula, 1900) (type species) from Iran and Sassanidotus gracilis thus zarudnyi sarghadensis Birula, 1903, syn. n.) from Afghanistan, Iran, and Pakistan. The neotype is designated for S. zarudnyi.


Scorpions Of Iran (Arachnida: Scorpiones). Part Vii. Kerman Province, Shahrokh Navidpour, Majeed Ezatkhah, František Kovařík, Michael E. Soleglad, Victor Fet Dec 2016

Scorpions Of Iran (Arachnida: Scorpiones). Part Vii. Kerman Province, Shahrokh Navidpour, Majeed Ezatkhah, František Kovařík, Michael E. Soleglad, Victor Fet

Victor Fet

Thirteen species of scorpions belonging to two families are reported from the Kerman Province of Iran. Of these, the species Compsobuthus kaftani Kovařík, 2003, Mesobuthus macmahoni (Pocock, 1900), Orthochirus farzanpayi (Vachon et Farzanpay, 1987), Polisius persicus Fet, Capes et Sissom, 2001, Sassanidotus gracilis (Birula, 1900), and Hemiscorpius lepturus Peters, 1861 are recorded from the province for the first time. Kerman Province contains type localities of six species of scorpions, of which Kraepelinia palpator (Birula, 1903) and Orthochirus gruberi Kovařík et Fet, 2006 are valid. Prionurus crassicauda orientalis Birula, 1900 is a synonym of Androctonus crassicauda (Olivier, 1807), Buthus eupeus kirmanensis ...


Scorpions Of Iran (Arachnida, Scorpiones). Part Ix. Hormozgan Province, With A Description Of Odontobuthus Tavighiae Sp. N. (Buthidae), Shahrokh Navidpour, Michael E. Soleglad, Victor Fet, František Kovařík Dec 2016

Scorpions Of Iran (Arachnida, Scorpiones). Part Ix. Hormozgan Province, With A Description Of Odontobuthus Tavighiae Sp. N. (Buthidae), Shahrokh Navidpour, Michael E. Soleglad, Victor Fet, František Kovařík

Victor Fet

Twenty species of scorpions belonging to three families are reported from the Hormozgan Province of Iran. Of these, eight species and subspecies are recorded from the province for the first time: Buthacus macrocentrus (Ehrenberg, 1828), Compsobuthus persicus Navidpour et al., 2008, Iranobuthus krali Kovařík, 1997, Mesobuthus eupeus persicus (Pocock, 1899), Mesobuthus phillipsii (Pocock, 1889), Odontobuthus bidentatus Lourenço & Pézier, 2002, Odontobuthus doriae (Thorell, 1876), and Razianus zarudnyi (Birula, 1903). Odontobuthus tavighiae sp. n. is described and compared with all species of the genus Odontobuthus Vachon, 1950. Paraorthochirus goyffoni Lourenco et Vachon, 1995 is synonymized with Orthochirus farzanpayi (Vachon et Farzanpay, 1987 ...


Scorpions Of Iran (Arachnida, Scorpiones). Part V. Chahar Mahal & Bakhtiyari Province, Khodadad Pirali-Kheirabadi, Shahrokh Navidpour, Victor Fet, František Kovařík, Michael E. Soleglad Dec 2016

Scorpions Of Iran (Arachnida, Scorpiones). Part V. Chahar Mahal & Bakhtiyari Province, Khodadad Pirali-Kheirabadi, Shahrokh Navidpour, Victor Fet, František Kovařík, Michael E. Soleglad

Victor Fet

Ten species of scorpions belonging to three families are reported from the montane Chahar Mahal & Bakhtiyari Province of Iran. Of these, only Razianus zarudnyi (Birula, 1903) was formerly reported for this province, while nine species are reported for the first time: Androctonus crassicauda (Olivier, 1807), Compsobuthus matthiesseni (Birula, 1905), Hottentotta saulcyi (Simon, 1880), Hottentotta zagrosensis Kovařík, 1997, Mesobuthus eupeus phillipsii (Pocock, 1889), Odontobuthus doriae (Thorell, 1876), Orthochirus zagrosensis Kovařík, 2004, Hemi- scorpius lepturus Peters, 1861, and Scorpio maurus townsendi (Pocock, 1900). Also presented is a key to all species of scorpions found in the Chahar Mahal & Bakhtiyari Province.


Scorpions Of Iran (Arachnida: Scorpiones). Part Vi. Lorestan Province, Shahrokh Navidpour, Hassan H. Nayebzadeh, Michael E. Soleglad, Victor Fet, František Kovařík, Mohammad Hassan Kayedi Dec 2016

Scorpions Of Iran (Arachnida: Scorpiones). Part Vi. Lorestan Province, Shahrokh Navidpour, Hassan H. Nayebzadeh, Michael E. Soleglad, Victor Fet, František Kovařík, Mohammad Hassan Kayedi

Victor Fet

Ten species of scorpions belonging to three families are reported from the Lorestan Province of Iran. Of these, five species are recorded from the province for the first time: Hottentotta zagrosensis Kovařík, 1997; Mesobuthus eupeus phillipsii (Pocock, 1889); Orthochirus iranus Kovařík, 2004; Razianus zarudnyi (Birula, 1903) ; and Scorpio maurus townsendi (Pocock, 1900). One new species is described, Hottentotta lorestanus sp. n.; it can be easily distinguished from the other four species of the genus known from Iran by its coloration; it is the only Iranian species which has the entire pedipalps yellow and the metasomal segments I to IV greenish ...


Scorpions Of Iran (Arachnida: Scorpiones). Part Vi. Lorestan Province, Shahrokh Navidpour, Hassan H. Nayebzadeh, Michael E. Soleglad, Victor Fet, František Kovařík, Mohammad Hassan Kayedi Dec 2016

Scorpions Of Iran (Arachnida: Scorpiones). Part Vi. Lorestan Province, Shahrokh Navidpour, Hassan H. Nayebzadeh, Michael E. Soleglad, Victor Fet, František Kovařík, Mohammad Hassan Kayedi

Victor Fet

Ten species of scorpions belonging to three families are reported from the Lorestan Province of Iran. Of these, five species are recorded from the province for the first time: Hottentotta zagrosensis Kovařík, 1997; Mesobuthus eupeus phillipsii (Pocock, 1889); Orthochirus iranus Kovařík, 2004; Razianus zarudnyi (Birula, 1903); and Scorpio maurus townsendi (Pocock, 1900). One new species is described, Hottentotta lorestanus sp. n.; it can be easily distinguished from the other four species of the genus known from Iran by its coloration; it is the only Iranian species which has the entire pedipalps yellow and the metasomal segments I to IV greenish ...


Scorpions Of Iran (Arachnida, Scorpiones). Part Iv. Kohgilouyeh & Boyer Ahmad Province, Shahrokh Navidpour, František Kovařík, Michael E. Soleglad, Victor Fet Dec 2016

Scorpions Of Iran (Arachnida, Scorpiones). Part Iv. Kohgilouyeh & Boyer Ahmad Province, Shahrokh Navidpour, František Kovařík, Michael E. Soleglad, Victor Fet

Victor Fet

According to previous surveys, only Mesobuthus eupeus phillipsii (Pocock, 1889), Orthochirus zagrosensis Kovařík, 2004, and Hemiscorpius lepturus Peters, 1861 have been recorded from Kohgilouyeh & Boyer Ahmad Province. Collections made by a team under Shahrokh Navidpour (Razi Reference Laboratory of Scorpion Research, Razi Vaccine and Serum Research Institute, Ahvaz, Khoozestan, Iran) reveal eight other species belonging to two families recorded from the province for the first time: Androctonus crassicauda (Olivier, 1807); Compsobuthus matthiesseni (Birula, 1905); Hottentotta saulcyi (Simon, 1880); Hottentotta zagrosensis Kovařík, 1997; Odontobuthus bidentatus Lourenço et Pézier, 2002; Orthochirus iranus Kovařík, 2004; Razianus zarudnyi (Birula, 1903); and Scorpio maurus townsendi (Pocock, 1900). Also presented is a key to all species of scorpions found in the province.


Scorpions Of Iran (Arachnida, Scorpiones). Part Viii. Fars Province, Shahrokh Navidpour, Victor Fet, František Kovařík, Michael E. Soleglad Dec 2016

Scorpions Of Iran (Arachnida, Scorpiones). Part Viii. Fars Province, Shahrokh Navidpour, Victor Fet, František Kovařík, Michael E. Soleglad

Victor Fet

18 species of scorpions belonging to three families are reported from the Fars Province of Iran. Of these, four species are recorded from the province for the first time: Compsobuthus persicus Navidpour, Soleglad, Fet et Kovařík, 2008, Mesobuthus eupeus persicus (Pocock, 1899), Odontobuthus bidentatus Lourenço et Pézier, 2002, and Scorpio maurus townsendi (Pocock, 1900). Also presented is a key to all species of scorpions found in the province.


Scorpions Of Iran (Arachnida, Scorpiones). Part Iii. Ilam Province, Shahrokh Navidpour, Victor Fet, František Kovařík, Michael E. Soleglad Dec 2016

Scorpions Of Iran (Arachnida, Scorpiones). Part Iii. Ilam Province, Shahrokh Navidpour, Victor Fet, František Kovařík, Michael E. Soleglad

Victor Fet

Seven species of scorpions belonging to three families have been previously recorded from the Ilam Province of Iran: Androctonus crassicauda (Olivier, 1807), Compsobuthus matthiesseni (Birula, 1905), Hottentotta saulcyi (Simon, 1880), Mesobuthus eupeus phillipsii (Pocock, 1889), Razianus zarudnyi (Birula, 1903), Scorpio maurus townsendi (Pocock, 1900), and Hemiscorpius lepturus Peters, 1861. Collections made by a team organized by Shahrokh Navidpour (Razi Reference Laboratory of Scorpion Research, Razi Vaccine and Serum Research Institute, Ahvaz, Khoozestan, Iran) reveal seven other species recorded from the province for the first time: Apistobuthus susanae Lourenço, 1998, Buthacus macrocentrus (Ehrenberg, 1828), Compsobuthus jakesi Kovařík, 2003, Odontobuthus bidentatus Lourenço ...


Scorpions Of Iran (Arachnida, Scorpiones). Part Ii. Bushehr Province, Shahrokh Navidpour, Michael E. Soleglad, Victor Fet, František Kovařík Dec 2016

Scorpions Of Iran (Arachnida, Scorpiones). Part Ii. Bushehr Province, Shahrokh Navidpour, Michael E. Soleglad, Victor Fet, František Kovařík

Victor Fet

The Bushehr Province of Iran contains the type localities of three scorpion taxa, Mesobuthus eupeus phillipsii (Pocock, 1889), Orthochirus iranus Kovařík, 2004, and Scorpio maurus townsendi (Pocock, 1900). Apart from them, Androctonus crassicauda (Olivier, 1807), Compsobuthus matthiesseni (Birula, 1905), Hottentotta saulcyi (Simon, 1880), Odontobuthus bidentatus Lourenço & Pézier, 2002, Razianus zarudnyi (Birula, 1903), and Hemiscorpius lepturus Peters, 1862 have also been known to occur in the province. Collections made by a team under Shahrokh Navidpour (Razi Reference Laboratory of Scorpion Research, Razi Vaccine and Serum Research Institute, Ahvaz, Khoozestan, Iran) reveal four additional species recorded from the province for the first ...


Scorpions Of Iran (Arachnida, Scorpiones). Part I. Khoozestan Province, Shahrokh Navidpour, František Kovařík, Michael E. Soleglad, Victor Fet Dec 2016

Scorpions Of Iran (Arachnida, Scorpiones). Part I. Khoozestan Province, Shahrokh Navidpour, František Kovařík, Michael E. Soleglad, Victor Fet

Victor Fet

Collections made by a team of Shahrokh Navidpour (Razi Reference Laboratory of Scorpion Research, Razi Vaccine and Serum Research Institute, Ahvaz, Khoozestan, Iran) include 17 of the 19 species known to inhabit Khoozestan Province, and form the basis of this paper. Among them are two new species (Hottentotta khoozestanus sp. n. and Vachoniolus iranus sp. n.), Compsobuthus jakesi Kovařík, 2003 previously known only from Iraq, and five species representing first records for the province: Buthacus macrocentrus (Ehrenberg, 1828); Odontobuthus bidentatus Lourenço & Pézier, 2002; Orthochirus farzanpayi (Vachon et Farzanpay, 1987); Orthochirus stockwelli (Lourenço et Vachon, 1995) comb. n.; and Scorpio maurus townsendi (Pocock ...


Occurrence Of A Telson Gland In The Genus Superstitionia Stahnke, 1940 (Scorpiones: Superstitioniidae), Rolando Teruel, Graeme Lowe, Victor Fet, Kelly Anne Daniel Dec 2016

Occurrence Of A Telson Gland In The Genus Superstitionia Stahnke, 1940 (Scorpiones: Superstitioniidae), Rolando Teruel, Graeme Lowe, Victor Fet, Kelly Anne Daniel

Victor Fet

In this note, we record for the first time the occurrence of a telson gland in the monotypic genus Superstitionia Stahnke 1940 (Scorpiones: Superstitioniidae). The putative gland is marked by an elongated patch of wrinkled cuticle restricted to the dorso-apical surface of the telson vesicle of adult males, and is absent from females and immatures. Similar presumed glandular structures have been observed in a number of other scorpions, including many South American Bothriuridae, and the recently defined vaejovid genera Chihuahuanus González-Santillán et Prendini, 2013 and Maaykuyak González-Santillán et Prendini, 2013.


Laterobasal Aculear Serrations (Las) In Scorpion Family Vaejovidae (Scorpiones: Chactoidea), Victor Fet, Michael E. Soleglad, Michael S. Brewer Dec 2016

Laterobasal Aculear Serrations (Las) In Scorpion Family Vaejovidae (Scorpiones: Chactoidea), Victor Fet, Michael E. Soleglad, Michael S. Brewer

Victor Fet

The discovery of a new structure on the laterobasal aspect of the telson aculeus is described here for the first time. This structure, termed the laterobasal aculear serrations (LAS), is a row of minute denticles located on each side of the aculeus base, found exclusively in scorpion family Vaejovidae, thus potentially providing a new synapomorphy for this New World family. The LAS structure of major representative vaejovid genera and Vaejovis groups is illustrated with SEM micrographs. Also provided is a comprehensive list of non-vaejovid Recent scorpion genera examined where the LAS structure was found to be absent.


Etudes On Iurids, Iv. Observations On Calchas Gruberi From Megisti Island, Greece (Scorpiones: Iuridae), Iasmi Stathi, Victor Fet, Michael E. Soleglad Dec 2016

Etudes On Iurids, Iv. Observations On Calchas Gruberi From Megisti Island, Greece (Scorpiones: Iuridae), Iasmi Stathi, Victor Fet, Michael E. Soleglad

Victor Fet

A series of Calchas specimens from the Greek island of Megisti (= Kastelorizo) was examined. It is shown by detailed analysis of several key diagnostic characters that this population from Megisti Island belongs to C. gruberi Fet, Soleglad et Kovařík, 2009. Therefore, C. nordmanni Birula, 1899 is not present in Greek fauna. The population of C. gruberi from Megisti comprises the largest specimens so far reported.


High-Level Systematics And Phylogeny Of The Extant Scorpions (Scorpiones: Orthosterni), Michael E. Soleglad, Victor Fet Dec 2016

High-Level Systematics And Phylogeny Of The Extant Scorpions (Scorpiones: Orthosterni), Michael E. Soleglad, Victor Fet

Victor Fet

A number of authors (e. g. Birula, 1917a, 1917b; Mello-Leitão, 1945; Stockwell, 1989) addressed above-level systematics of extant scorpions, and accepted the grouping of scorpion families in several superfamilies. At the same time, Kjellesvig-Waering (1986) classified all extant scorpions under the same superfamily, Scorpionoidea. Sissom (1990) and Fet et al. (2000) did not list any superfamilies, considering the systematic situation above family (and often at the family level as well) unresolved. Most recently, Lourenço (2000a) listed six superfamilies, largely following the unpublished but important study of Stockwell (1989). The goal of this paper is to address scorpion systematics and phylogeny ...